Publications

2008

  • Descamps, S., Boutin, S., Berteaux, D., McAdam, A.G., & Gaillard, J.-M. (2008) Cohort effects in red squirrels: the influence of density, food abundance and temperature on future survival and reproductive success. Journal of Animal Ecology 77: 305-314.
  • Lane, J.E., Boutin, S., Gunn, M.R., Slate, J., & Coltman, D.W. (2008) Female multiple mating and paternity in free-ranging North Aerican red squirrels. Animal Behaviour 75: 1927-1937.
  • Laura Guichon, M. & Patrick Doncaster, C. Invasion dynamics of an introduced squirrel in Argentina. Ecography 0, 0-0. OnlineEarly Article
  • Perry, D.A.N. & Perry, G.A.D. (2008) Improving interactions between animal rights groups and conservation biologists. Conservation Biology 22: 27-35.
  • Steele, M.A., Halkin, S.L., Smallwood, P.D., McKenna, T.J., Mitsopoulos, K., & Beam, M. (2008) Cache protection strategies of a scatter-hoarding rodent: do tree squirrels engage in behavioural deception. Animal Behaviour 75, 705-714.
  • Wauters, L.A, Githiru, M., Bertolino, S., Molinari, A., Tosi, G., & Lens, L. Demography of alpine red squirrel populations in relation to fluctuations in seed crop size. Ecography 31, 140-160./li>

2007

  • Boon, A.K., Reale, D., & Boutin, S. (2007) The interaction between personality, offspring fitness and food abundance in North American red squirrels. Ecology Letters 10: 1094-1104.
  • Descamps, S., Boutin, S., Berteaux, D., & Gaillard, J.-M. (2007) Female red squirrels fit Williams' hypothesis of increasing reproductive effort with increasing age. Journal of Animal Ecology, 76: 1192-1201.
  • Duff, J., Higgins, R., & Farrelly, S. (2007) Enteric adenovirus infection in a red squirrel (Sciurus vulgaris). The Veterinary Record March 17, 2007: 384.
  • Edelman, A.J. & Koprowski, J.L. (2007) Communal nesting in asocial Abert's squirrels: the role of social thermoregulation and breeding strategy. Ethology 113: 147-154.
  • Finnegan, L.A., Edwards, C.J. & Rochford, J.M. (2007) Origin of, and conservation units in, the Irish red squirrel (Sciurus vulgaris) population. Conservation Genetics DOI 10.1007/s10592-007-9419-7.
  • Finnegan, L., Hamilton, G., Perol, J. & J Rochford, J. (2007) The use of hair tubes as an indirect method for monitoring red and grey squirrel populations. Biology and the Environment 107B: 55-60.
  • Hurme, E., Reunanen, P., Monkkonen, M., Nikula, A., Nivala, V., & Oksanen, J. (2007) Local habitat patch pattern of the Siberian flying squirrel in a managed boreal forest landscape. Ecography 30: 277-287.
  • Lane, J., Boutin, S., Slate, J., & Coltman, D. (2007) Genetic relatedness of mates does not predict patterns of parentage in North American red squirrels. Animal Behaviour74: 611-619.
  • Lawton, C. & Rochford, J. (2007) The recovery of grey squirrel (Sciurus carolinensis) populations after intensive control programmes. Biology and Environment: Proceedings of the Royal Irish Academy 107B: 19-29.
  • Selonen, V., Hanski, & Desrochers, A. (2007) Natal habitat-biased dispersal in the Siberian flying squirrel. Proceedings of the Royal Society London B 274: 2063-2068.
  • Siepielski, A.M. & Benkman, C.W. (2007) Selection by a predispersal seed predator constrains the evolution of avian seed dispersal in pines. Functional Ecology 21: 611-618.
  • Tamura, N. & Hayashi, F. (2007) Five-year study of the genetic structure and demography of two subpopulations of the Japanese squirrel (Sciurus lis) in a continuous forest and an isolated woodlot. Ecological Research 22: 261-267.
  • Wauters, L., Preatoni, D., Molinari, A., & Tosi, G. (2007) Radio-tracking squirrels: performance of home range density and linkage estimators with small range and sample size. Ecological Modelling 202: 333-344.
  • Wauters, L.A., Vermeulen, M., Van Dongen, S., Bertolino, S., Molinari, A., Tosi, G., & Matthysen, E. (2007) Effects of spatio-temporal variation in food supply on red squirrel Sciurus vulgaris body size and body mass and its consequences for some fitness components. Ecography 30: 51-65.

2006

  • Boutin, S., Wauters, L., McAdam, A., Humphries, M., Tosi, G., & Dhondt, A. (2006) Anticipatory reproduction and population growth in seed predators. Science 314, 1928 - 1930.
  • Carvalho, L.d.S., Cowing, J., Wilkie, S., Bowmaker, J., & Hunt, D. (2006) Shortwave visual sensitivity in tree and flying squirrels reflects changes in lifestyle. Current Biology 16: R81-R83.
  • Edelman, A.J. & Koprowski, J.L. (2006) Diet and tree use of Abert’s squirrels (Sciurus aberti) in a mixed-conifer forest. Southwestern Naturalist 50: 461-465.
  • Edelman, A.J. & Koprowski, J.L. (2007) Communal Nesting in Asocial Abert's Squirrels: the Role of Social Thermoregulation and Breeding Strategy. Ethology 113, 147-154.
  • Ferryman, M., Mayle, B., & Morgan, G. (2006) Visual methods for evaluating the state of sexual development in male grey squirrels (Sciurus carolinensis). Reproduction, Fertility and Development 18, 283-293.
  • Gurnell, J., Rushton S.P., Lurz, P.W.W., Sainsbury, A.W., Nettleton, P, Shirley, M.D.F., Bruemmer, C. & Geddes, N. (2006) Squirrel poxvirus; landscape scale strategies for managing disease threat. Biological Conservation 131: 287-295.
  • Jansen, P.A., Bongers, F., & H. T. Prins, H. (2006) Tropical rodents change rapidly germinating seeds into long-term food supplies. Oikos 113, 449-458.
  • Meijaard, E., Kitchener, A.C., & Smeenk, C. (2006) 'New Bornean carnivore' is most likely a little known flying squirrel. Mammal Review 36, 318-324.
  • McInnes, C.J., Wood, A.R., Thomas, K., Sainsbury, A.W., Gurnell, J., Dein, F.J. and Nettleton. P.F. (2006) Genomic characterisation of a novel poxvirus contributing to the decline of the red squirrel (Sciurus vulgaris) in the UK. Journal General Virology 87: 2115-2125.
  • Molinari, A., Wauters L.A., Airoldi, G., Cerinotti, F., Martinoli, A. & Tosi, G.(2006) Cone selection by Eurasian red squirrels in mixed conifer forests in the Italian Alps.Acta Oecologica 30: 1-10. (doi:10.1016/j.actao.2005.11.004).
  • Moore, J. & Swihart, R. (2006) Modeling patch occupancy by forest rodents: incorporating detectability and spatial autocorrelation with hierarchically structured data. Journal of Wildlife Management 69, 933-949.
  • Ozgul, A., Armitage, K.B., Blumstein, D.T., Vanvuren, D.H., & Oli, M.K. (2006) Effects of patch quality and network structure on patch occupancy dynamics of a yellow-bellied marmot metapopulation. Journal of Animal Ecology 75, 191-202.
  • Robitaille, J.-F. & Linley, R.D. (2006) Structure of forests used by small mammals in the industrially damaged landscape of Sudbury, Ontario, Canada. Forest Ecology and Management 225: 160-167.
  • Rushton S.P., Lurz, P.W.W., Gurnell, J., Nettleton, P. , Bruemmer, C.,Shirley, M.D.F. and Sainsbury, A.W. (2006) Disease threats posed by alien species: the role of a poxvirus in the decline of the native red squirrel in Britain. Epidemiology and Infection 134: 521-533.
  • Selonen,V. & Hanski,I.K. (2006) Habitat exploration and use in dispersing juvenile flying squirrels. Journal of Animal Ecology 75:1440-1449.
  • Steele, M.A., Manierre, S., Genna, T., Contreras, T.A., Smallwood, P.D., & Pereira, M.E. (2006) The innate basis of food-hoarding decisions in grey squirrels: evidence for behavioural adaptations to the oaks. Animal Behaviour 71: 155-160.
  • Sushma, H.S. & Singh, M. (2006) Resource partitioning and interspecific interactions among sympatric rain forest arboreal mammals of the Western Ghats, India. Behavioral Ecology 17: 479-490.
  • Tattoni, C., Preatoni D.G., Lurz, P.W.W., Rushton, S.P., Tosi, G., Bertolino, S., Martinoli, A., & Wauters, L.A. (2006) Modelling the expansion of a grey squirrel population: implications for squirrel control. Biological Invasions8: 1605-1619.
  • Thorington Jr, R. & Ferrell, K. (2006) Squirrels: The Animal Answer Guide. 208 pp. John Hopkins University Press, Baltimore, USA. Link to JHUP.
  • Wagner, D.M., Drickamer, L.C., Krpata, D.M., Allender, C.J., Van Pelt, W.E., & Keim, P. (2006) Persistence of Gunnison's prairie dog colonies in Arizona, USA. Biological Conservation 130, 331-339.
  • Winterrowd, M.F. & Weigl, P.D. (2006) Mechanisms of Cache Retrieval in the Group Nesting Southern Flying Squirrel (Glaucomys volans). Ethology 112, 1136-1144.
  • Xiao, Z., Jansen, P.A., & Zhang, Z. (2006) Using seed-tagging methods for assessing post-dispersal seed fate in rodent-dispersed trees. Forest Ecology and Management 223: 18-23.

2005

  • Bryce, J., Cartmel, S. & Quinn, C.P. (2005) Habitat use by red and grey squirrels: results of two recent studies and implications for management. Information Note. Forestry Commission, Edinburgh. 12pp.
  • Edelman, A.J. & Koprowski, J.L. (2005) Selection of drey sites by Abert’s squirrels in an introduced population. Journal of Mammalogy, 86(6):1220-1226.
  • Edelman, A.J., Koprowski, J.L. & Edelman, J.R. (2005) Kleptoparasitic behavior and species richness at Mt. Graham red squirrel middens. In Gottfried, Gerald J.; Gebow, Brooke S.; Eskew, Lane G.; and Edminster, Carleton B., compilers. Connecting mountain islands and desert seas: biodiversity and management of the Madrean Archipelago II. 2004 May 11-15; Tucson, AZ. Proceedings RMRS-P-36. pp 395-398. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station. 631 p.
  • Gerhardt, F. (2005) Food pilfering in larder-hoarding red squirrels (Tamiascsiurus hudsonicus). Journal of Mammalogy 86: 108-114.
  • Goheen, J. & Swihart, R. (2005) Resource selection and predation of North American red squirrels in deciduous forest fragments. Journal of Mammalogy 86: 22-28.
  • Hurme, E., Monkkonen, M., Nikula, A.,Nivala, V., Reunanen, P., Heikkinen, T., & Ukkola, M. (2005) Building and evaluating predictive occupancy models for the Siberian flying squirrel using forest planning data. Forest Ecology and Management 216:241-256.
  • Koprowski, J.L. (2005) Management and conservation of tree squirrels: the importance of endemism, species richness, and forest condition. In Gottfried, Gerald J.; Gebow, Brooke S.; Eskew, Lane G.; and Edminster, Carleton B., compilers. Connecting mountain islands and desert seas: biodiversity and management of the Madrean Archipelago II. 2004 May 11-15; Tucson, AZ. Proceedings RMRS-P-36. pp. 245-250. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station. 631 p.
  • Koprowski, J.L. (2005) Annual cycles in body mass and reproduction of endangered Mt. Graham red squirrels. Journal of Mammalogy 86: 309-313.
  • Koprowski, J.L. (2005) The response of tree squirrels to fragmentation: a review and synthesis. Animal Conservation 8: 369-376.
  • Koprowski, J.L. (2005, July 20). Pine Squirrel (Tamiasciurus hudsonicus): a technical conservation assessment. [Online]. USDA Forest Service, Rocky Mountain Region. Available: http://www.fs.fed.us/r2/projects/scp/assessments/pinesquirrel.pdf [date of access].
  • Koprowski, J.L., Alanen, M.I. & Lynch, A.M.(2005) Nowhere to run and nowhere to hide: Response of endemic Mt. Graham red squirrels to catastrophic forest damage. Conservation Biology 126: 491-498.
  • Koprowski, J.L. & Corse, M.C. (2005) Time budgets, activity periods, and behavior of Mexican fox squirrels. Journal of Mammalogy 86: 947-952.
  • Koprowski, J.L. Edelman, A.J., Pasch, B.S. and Buecher, D.C. (2005) A dearth of data on the mammals of the Madrean Archipelago: What we think we know and what we actually do know. In Gottfried, Gerald J.; Gebow, Brooke S.; Eskew, Lane G.; and Edminster, Carleton B., compilers. Connecting mountain islands and desert seas: biodiversity and management of the Madrean Archipelago II. 2004 May 11-15; Tucson, AZ. Proceedings RMRS-P-36. pp 412-415. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station. 631 p.
  • Lawton, C. (Ed.) (2005). Proceedings of the Irish Red Squirrel Conservation Symposium , 22nd April, 2005. 20pp. National University of Ireland, Galway. (PDF)
  • Lu, J. & Zhang, Z. (2005) Food hoarding behaviour of David's rock squirrel Sciurotamias davidianus. Acta Zoologica Sinica 51: 376-382.
  • Mezquida, E.T. & Benkman, C.W. (2005) The geographic selection mosaic for squirrels, crossbills and Aleppo pine. Journal of Evolutionary Biology 18: 348-357.
  • Oshida, T., AramovV, A., Yanagawa, H., & Masuda, R. (2005) Phylogeography of the Russian flying squirrel (Pteromys volans): implication of refugia theory in arboreal small mammal of Eurasia. Molecular Ecology, 14, 1191-1196.
  • Pasch, B.S. & Koprowski, J.L. (2005) Correlates of vulnerability in Chiricahua fox squirrels. In Gottfried, Gerald J.; Gebow, Brooke S.; Eskew, Lane G.; and Edminster, Carleton B., compilers. Connecting mountain islands and desert seas: biodiversity and management of the Madrean Archipelago II. 2004 May 11-15; Tucson, AZ. Proceedings RMRS-P-36. pp 426-428. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station. 631 p.
  • Selonen, V., Painter, J. N. & Hanski, I. K. (2005) Microsatellite variation in the Siberian flying squirrel in Finland. Annales Zoologici Fennici 42: 505:511.
  • Steele, M., Wauters, L.A. & Larsen, K.A. (2005) Selection, Predation and Dispersal of Seeds by Tree Squirrels in Temperate and Boreal Forests: are Tree Squirrels Keystone Granivores? In Seed Fate (Eds. P.-M. Forget, J.E. Lambert, P.E. Hulme and S.B. Vander Wall). pp 205-221. CAB International.
  • Trizio, I., B. Crestanello,B., Galbusera, P., Wauters, L.A., Tosi, G. Matthysen, E. & Hauffe, H. (2005) Geographical distance and physical barriers shape the genetic structure of Eurasian red squirrels (Sciurus vulgaris) in the Italian Alps. Molecular Ecology 14: 469:481.
  • Wauters,L.A., Bertolino, S., Adamo, M., Van Dongen, S., Tosi, G. (2005) Food shortage disrupts social organization: the case of red squirrels in conifer forests. Evolutionary Ecology 19:375-404
  • Wauters, L., Tosi, G., & Gurnell, J. (2005) A review of the competitive effects of alien grey squirrels on behaviour, activity and habitat use of red squirrels in mixed, deciduous woodland in Italy. Hystrix It. J. Mamm. 16: 27-40.

2004

  • Bakker, V.J.& Van Vuren, D.H. (2004) Gap-crossing decisions by the Red Squirrel, a forest-dependent small mammal. Conservation Biology 18: 689-697.
  • Gurnell, J. (2004) Grey Squirrels Invade Europe. Mammals UK. Mammals Trust UK, London
  • Gurnell,J., Lurz, P.W.W., Shirley, M.D.F., Cartmel, S., Garson, P.J., Magris, L., & Steele, J. (2004) Monitoring red (Sciurus vulgaris) and grey (Sciurus carolinensis) squirrels in Britain. Mammal Review 34: 51-74.
  • Gurnell, J., Wauters, L.A., Lurz, P.W.W. & Tosi, G. (2004) Alien species and interspecific competition: effects of introduced eastern grey squirrels on red squirrel population dynamics. Journal of Animal Ecology 73: 26-35.
  • Hale, M. L., Lurz, P. W. W. & Wolff, K. (2004). Patterns of genetic diversity in the red squirrel: footprints of biogeographic history and artificial introductions. Conservation Genetics 5:167-179.
  • Haughland, D. & Larsen, K. (2004) Ecology of North American red squirrels across contrasting habitats: relating natal dispersal to habitat.Journal of Mammalogy 85: 225-236.
  • Haughland, D.L. & Larsen, K.W. (2004) Exploration correlates with settlement: red squirrel dispersal in contrasting habitats. Journal of Animal Ecology 73: 1024-1034.
  • Hewson, C., Fuller, R., Mayle, B. & Smith, K. (2004) Possible impacts of grey squirrels on birds and other wildlife. British Wildlife 15: 183-191.
  • Lehmkuhl, J.F. Gould, L.E.,Cázaresc, E & Hosford, D.R. (2004) Truffle abundance and mycophagy by northern flying squirrels in eastern Washington forests. Forest Ecology and Management 200: 49-65.
  • Linders, M., West, S.& Vander Haegen, W.M. (2004) Seasonal variability in the use of space by western gray squirrels in Southcentral Washington. Journal of Mammalogy 85: 511-516.
  • Mayle, B. & Gurnell, J. (2004) Increasing efficiency of grey squirrel control in conifer habitats. Forestry and British Timber April 2004.
  • Mayle, B. & Gurnell, J. (2004) Comparison of trap location and bait type for grey squirrel control in conifers. 20 pp. Report to the Sustainable Forestry Group, Forestry Commission, Edinburgh.
  • Miyamoto, A., Tamura, N., Sugimura, K. & Yamada, F.(2004) Predicting habitat distribution of the alien Formosan squirrels using logistic regression model. Global Environmental Research 8: 13-21.
  • Painter, J., Selonen, V. & Hanski, I. K. (2004) Microsatellite loci for the Siberian flying squirrel, Pteromys volans. Molecular Ecology Notes 4:119-121.
  • Sainsbury, A., Kountouri, A., DuBoulay, G., & Kertesz, P. (2004) Oral disease in free-living red squirrels (Sciurus vulgaris) in the United Kingdom.Journal of Wildlife Diseases, 40, 185-196.
  • Selonen, V. & Hanski, I. K. (2004) Young flying squirrels (Pteromys volans)dispersing in fragmented forests. Behavioral Ecology 15: 564-571.
  • Tamura, N. (2004) Effects of habitat mosaic on home range size of the Japanese squirrel Sciurus lis." Mammal Study 29: 9-14.
  • Wauters L.A., Zaninetti M., Tosi G., Bertolino S.(2004) Is coat-colour polymorphism in Eurasian red squirrels (Sciurus vulgaris L.) adaptive? Mammalia 68:37-48.
  • Wauters, L.A., Matthysen, E. Adriaensen, F. & Tosi, G. (2004) Within-sex density-dependence and population dynamics of red squirrels Sciurus vulgaris Journal of Animal Ecology 73:11-25.
  • Wells, K., Pfeiffer, M., Lakim, M.B. & Linsenmair, K.E. (2004) Use of arboreal and terrestrial space by a small mammal community in a tropical rain forest in Borneo, Malaysia. Journal of Biogeography 31: 641-652.

2003

  • Anon (2003)Urban grey squirrels. DEFRA Rural Develeopment Setrvices Technical Advisory Note WM09 [PDF]
  • Bertolino, S. Wauters, L.A., De Bruyn L. & Canestri-Trotti, G. (2003) Prevalence of coccidia parasites (Protozoa) in red squirrels ( Sciurus vulgaris): effects of host phenotype and environmental factors. Oecologia 137, 286-295.
  • Desrochers, A., Hanski, I. K. & Selonen, V. (2003) Siberian flying squirrel responses to high- and low- contrast forest edges. Landscape Ecology 18:543-552.
  • Goheen, J. & Swihart, R. (2003) Food-hoarding behavior of gray squirrels and North American red squirrels in the central hardwoods region: implications for forest regeneration. Canadian Journal of Zoology, 81, 1636-1639.
  • Goheen, J., Swihart, R., Gehring, T., & Miller, M. (2003) Forces structuring tree squirrel communities in landscapes fragmented by agriculture: species differences in perceptions of forest connectivity and carrying capacity. Oikos, 102, 95-103.
  • Goheen, J., Swihart, R., & Robins, J. (2003) The anatomy of a range expansion: changes in cranial morphology and rates of energy extraction for North American red squirrels from different latitudes. Oikos, 102, 33-44.
  • Hale, M. L. & Lurz, P. W. W. (2003). Morphological changes in a British mammal as a result of introductions and changes in landscape management: the red squirrel (Sciurus vulgaris). Journal of Zoology, London: 260: 159-167.
  • Hutton KA, Koprowski JL, Greer VL, Alanen MI, Schauffert CA & Young PJ. (2003) Use of mixed-conifer and spruce-fir forests by an introduced population of Abert's squirrels (Sciurus aberti). Southwestern Naturalist 48:257-260.
  • Lurz, P. W. W., Geddes, N., Lloyd, A. J., Shirley, M. D. F., Rushton, S. P. & Burlton, B. (2003) Planning a red squirrel conservation area: using a spatially explicit population dynamics model to predict the impact of felling and forest design plans. Forestry, 76, 95-108.
  • Lurz, P. W. W., Rushton, S. P. & Gurnell. J. (2003) Kielder Red Squirrel Project. Report for FE Kielder Forest and Mammals Trust UK. 22 pp.
  • Petty, S. J., Lurz, P. W. W. & Rushton, S. P. (2003) Predation of red squirrels by northern goshawks in a conifer forest in northern England: can this limit squirrel numbers and create a conservation dilemma? Biological Conservation, 111, 105-114.
  • Réale, D., McAdam, A., Boutin, S. & Berteaux, D. (2003) Genetic and plastic response of a northern mammal to climate change. Proceedings of the Royal Society of London B 270: 591-596.
  • Réale, D., Berteaux, D., McAdam, A. & Boutin, S. (2003) Lifetime selection on heritable life-history traits in a natural population of red squirrels. Evolution 57: 2416-2423.
  • Selonen, V. & Hanski, I. K. (2003) Movements of the flying squirrel Pteromys volans in corridors and in matrix habitat. Ecography 26:641-651.
  • Stapleford, D. (2003) An Affair with Red Squirrels The Larks Press, Ordnance Farmhouse, Guist Bottom, Dereham, Norfolk NR20 SPF UK. Price £5. 58 pp.
  • Swihart, R., Atwood, T., Goheen, J., Scheiman, D., Munroe, K., & Gehring, T. (2003) Patch occupancy of North American mammals: is patchiness in the eye of the beholder? Journal of Biogeography, 30, 1259-1279.
  • Tompkins, D., Sainsbury, A.W., Netteleton, P., Buxton, D. & Gurnell. J. (2002) Parapoxvirus causes a deleterious disease of red squirrels associated with UK population declines. Proceedings of the Royal Society of London 269: 529-533.
  • Verbeylen, G., De Bruyn, L. & Matthysen, E. (2003) Patch occupancy, population density and dynamics in a fragmented red squirrel Sciurus vulgaris population. Ecography 26, 118-128.

2002

  • Barr, J. J., Lurz, P. W. W., Shirley, M. D. F. & Rushton, S. P. (2002) Evaluation of immunocontraception as a publicly acceptable form of vertebrate pest species control: the introduced grey squirrel in Britain as an example. Environmental Management, 30, 342-351.
  • Bertolini,S; Genovesi,P (2002) Spread and attempted eradication of the grey squirrel (Sciurus carolinensis) in Italy, and consequences for the red squirrel (Sciurus vulgaris) in Eurasia. Biological Conservation 109, 351-358.
  • Bryce,J; Johnson,PJ; Macdonald,DM (2002) Can niche use in red and grey squirrels offer clues for their apparent coexistence? Journal of Applied Ecology 39, 875-887.
  • Gurnell, J., Cartmel, S., Lurz, P.W.W. & Rushton, S.P. (2002) Modelling the effects of forest composition and management on conserving red squirrels in Clocaenog Forest, North Wales. Unpublished Report for Forestry Commission and Countryside Council for Wales.
  • Gurnell, J. & Steele, J. (2002) Grey Squirrel Control for Red Squirrel Conservation - Thetford Forest. Published Report for English Nature and the Forestry Commission.
  • Gurnell, J., Clark, M.J., Lurz, P.W.W., Shirley, M.D.F. & Rushton, S.P. (2002) Conserving red squirrels (Sciurus vulgaris): mapping and forecasting habitat suitability using a Geographic Information Systems Approach. Biological Conservation 105: 53-64.
  • Koprowski, J.L. (2002) Handling tree squirrels with an efficient and safe restraint. Wildlife Society Bulletin 30:101-103.
  • Kurtila,M; Pukkala,T; Loikkanen,J (2002) The performance of alternative spatial objective types in forest planning calculations: a case for flying squirrel and moose. Forest Ecology and Management 166, 245-260.
  • Lee,TH (2002) Feeding and hoarding behaviour of the Eurasian red squirrel Sciurus vulgaris during autumn in Hokkaido, Japan. Acta theriologica 147, 459-470.
  • Magris, L. & Gurnell, J. (2002) Population ecology of the red squirrel (Sciurus vulgaris) in a fragmented woodland ecosystem on the Island of Jersey, Channel islands. Journal of Zoology, London 256: 99-112.
  • McAdam,AG; Boutin,S; R‚ale,D; Berteaux,D (2002) Maternal effects and the potential for evolution in a natural population of animals. Evolution 56, 846-851.
  • Schauffert, C., J. Koprowski, V. Greer, M. Alanen & K. Hutton. (2002) Potential predators of Mt. Graham red squirrels. Southwestern Naturalist 47:498-501.
  • Wauters, L.A., Tosi, G. & Gurnell, J. (2002) Interspecific competition of grey on reds: do grey squirrels deplete tree seeds cached by red squirrels. Behavioral Ecology and Sociobiology 51:360-367
  • Tompkins, D., Sainsbury, A.W., Netteleton, P., Buxton, D. & Gurnell. J. (2002) Parapoxvirus causes a deleterious disease of red squirrels associated with UK population declines. Proceedings of the Royal Society of London 269: 529-533.

2001

  • Gurnell, J. (2001) Red squirrels: a species in decline. Mammals UK Summer 2001 Pp7-8, Mammals Trust UK, London.
  • Gurnell, J., Lurz, P.P.W. & Pepper, H. (2001) Practical techniques for surveying and monitoring squirrels. Forestry Commission Practice Note 11. Forestry Commission, Edinburgh. 12pp.
  • Gurnell, J. , Wauters, L.A., Preatoni, D. & Tosi, G. (2001) Spacing behaviour, kinship and dynamics of the grey squirrel in a newly colonised deciduous woodland in north Italy. Canadian Journal of Zoology 79: 1533-1543.
  • Hale, M. L., Peter W.W. Lurz, Mark D.F. Shirley, Steven Rushton, Robin M. Fuller & Kirsten Wolff (2001). The Impact of Landscape Management on the Genetic Structure of Red Squirrel Populations. Science 293: 2246-2248.
  • Koprowski, J.L. & M.C. Corse (2001) Food habits of Chiricahua Fox Squirrels (Sciurus nayaritensis chiricahuae). Southwestern Naturalist, 46:62-65.
  • Lurz, P.W.W., Rushton, S.P., Wauters, L.A., Bertolino, S., Corrado, I., Mazzoglio, P. & Shirley, M.D.F. (2001). Predicting gray squirrel expansion in North Italy: a spatially explicit modelling approach. Landscape Ecology , 16, 407-420
  • Sainsbury, A.W., Adair, B., Graham, D., Gurnell, J., Cunningham, A.A., Benko, M. & Papp, T. (2001) Isolation of a novel adenovirus associated with splenitis, diarrhoea, and mortality in translocated red squirrels, Sciurus vulgaris. Verh. ber. Ekrg. Zootiere 40: 265-270.
  • Selonen, V., Hanski, I. K & Stevens, P. (2001) Space use of Siberian flying squirrel Pteromys volans in fragmented forest landscapes. Ecography 24:588-600.
  • Shuttleworth, C.M. & Gurnell, J. (2001) The management of coastal sand dune woodland for red squirrels (Sciurus vulgaris L.). In Coastal Dune Management: Shared Experience of European Conservation Practice (eds. J.A. Houston, S.E. Edmondson & P.J. Rooney) pp. 117 - 127. Liverpool University Press, Liverpool, England.
  • Steele, M.A. & Koprowski, J.L. (2001) North American Tree Squirrels Smithsonian Institution Press, 201 pp.
  • Wauters, L.A., Gurnell, J., Martinoli, A. & Tosi, G. (2001) Does interspecific competition with grey squirrels affect the foraging behaviour and food choice of red squirrels. Animal Behaviour 61: 1079-1091.
  • Wauters, L.A., Gurnell, J. ,Preatoni, D. & Tosi, G (2001) Effects of spatial variation in food availability on spacing behaviour and demography of Eurasian red squirrels. Ecography 24: 525-538.

2000

  • Bruemmer, C., Lurz, P., Larsen, K. & Gurnell, J. (2000) Impacts and management of the alien Eastern Gray Squirrel in Great Britain and Italy: lessons for British Columbia. Pp. 341 - 349. In L. M. Darling (editor). Proceedings of a Conference on the Biology and Management of Species and Habitats at Risk, Kamloops, B.C., 15 - 19 Feb., 1999. Volume One. B.C. Ministry of Environment, Lands and Parks, Victoria, B.C. and University College of the Cariboo, Kamloops, B.C. 490 pp.
  • Cagnin, M., Aloise, G., Fiore, F., Oriolo, V. & Wauters, L.A. (2000). Habitat use and population densityof the red squirrel, Sciurus vulgaris meridionalis, in the Sila Grande mountain range (Calabria, South Italy). Ital. J. Zool. 67: 81-87.
  • Lurz, P.W.W., Garson, P.J. & Wauters, L.A.. (2000). Effects of temporal and spatial variations in food supply on the space and habitat use of red squirrels, Sciurus vulgaris L. J. Zool., Lond. 251: 167-178.
  • Rushton, S.P., Lurz, P.W.W. & Gurnell, J. (2000) Modelling the spatial dynamics of parapoxvirus disease in red and grey squirrels: a possible cause of the decline in the red squirrel in the United Kingdom? Journal of Applied Ecology 37:1-18.
  • Sainsbury, A.W., Nettleton, P., Gilray, J. & Gurnell, J. (2000) Grey squirrels have high seroprevalence to a parapoxvirus associated with deaths in red squirrels. Animal Conservation 3: 229-233.
  • Verbeylen, G. & De Bruyn, L. (2000). Inventarisatie van de Aziatische grondeekhoorn in De Panne. Een project van het IBW en AMINAL afdeling Natuur. Rapport 3 - September 2000.
  • Wauters, L.A. (2000). Squirrels - Medium-sized Granivores in Woodland Habitats. - In: Halle, S. and Stenseth N. C. (eds), Activity Patterns in Small Mammals: a Comparative Ecological Approach. Ecological Studies 141, Springer-Verlag, Heidelberg, Germany, pp. 131-143.
  • Wauters, L.A., Lurz, P.W.W. & Gurnell, J. (2000). The effects of interspecific competition by grey squirrels (Sciurus carolinensis) on the space use and population dynamics of red squirrels (S. vulgaris) in conifer plantations. Ecological Research 15: 271-284.
  • Teangana, D.Ó., Russ, J.M., Mathers, R.G., & Montgomery, W.I. (2000) Habitat associations of the red squirrel Sciurus vulgaris and grey squirrel S. carolinensis in Northern Ireland. Biology and Environment: Proceedings of the Royal Irish Academy, Vol. 100B, No. 1, 27-33.

1999

  • Barratt, E.M., Gurnell, J., Malarky, G. Deaville, R. & Bruford, M.W. (1999) Genetic structure of fragmented populations of red squirrel (Sciurus vulgaris) in Britain. Molecular Ecology 8: S55-S63.
  • David-Gray, Z.K., Gurnell, J. & Hunt, D.M. (1999) Estimating the relatedness in a population of grey squirrels, Sciurus carolinensis, using DNA fingerprinting. Acta theriologica 44: 243-251.
  • David-Gray, Z.K., Gurnell, J. & Hunt, D.M. (1999) DNA fingerprinting reveals high levels of genetic diversity in British populations of the introduced non-native grey squirrel, Sciurus carolinensis. Journal of Zoology, London 246: 443-486.
  • Gurnell, J. (1999) Grey squirrels in woodlands: managing grey squirrels to prevent woodland damage. Enact 7: 10-14.
  • Gurnell, J. and Wauters, L. (1999) Sciurus vulgaris. In The Atlas of European Mammals. Mitchell-Jones, A.J., Amori, G., Bogdanowicz, W., Krystufek, B., Reijnders, P.J.H., Spitzenberger. F., Stubbe, M., Thissen, J.B.M., Vohohralik, V. & Zima, J. (Eds.) pp. 180-181. Academic press, London.
  • Gurnell, J. and Wauters, L. (1999) Sciurus carolinensis. In The Atlas of European Mammals. Mitchell-Jones, A.J., Amori, G., Bogdanowicz, W., Krystufek, B., Reijnders, P.J.H., Spitzenberger. F., Stubbe, M., Thissen, J.B.M., Vohohralik, V. & Zima, J. (Eds.) pp. 178-179. Academic press, London.
  • Gurnell, J. and Wauters, L. (1999) Callosciurus erythraeus. In The Atlas of European Mammals. Mitchell-Jones, A.J., Amori, G., Bogdanowicz, W., Krystufek, B., Reijnders, P.J.H., Spitzenberger. F., Stubbe, M., Thissen, J.B.M., Vohohralik, V. & Zima, J. (Eds.) pp. 182-183. Academic press, London.
  • Garvish, L. and Gurnell, J. (1999) Sciurus anomalus. In The Atlas of European Mammals. Mitchell-Jones, A.J., Amori, G., Bogdanowicz, W., Krystufek, B., Reijnders, P.J.H., Spitzenberger. F., Stubbe, M., Thissen, J.B.M., Vohohralik, V. & Zima, J. (Eds.) pp. 176-177. Academic press, London.
  • Massingham Hart, R. (1999) Squirrel Proofing your Home and Garden. Workman Publishing Company. ISBN: 1580171915
  • Mathias, M.L. & Gurnell, J. (1999) Status and conservation of the red squirrel (Sciurus vulgaris L) in Portugal. Hystrix 10: 13-19.
  • Osborn, D.A. (1999) Squirrel dog basics: a guide to hunting squirrels with dogs. Tree Top Publications, Watkinsville, Georgia, 151 pp.
  • Rushton, S.P., Lurz, P.W.W., South, A.B. & Mitchell-Jones, A. (1999). Modelling the distribution of red squirrels on the Isle of Wight. Animal Conservation, 2, 111-121
  • Verbeylen, G., De Bruyn, L. & Matthysen, E. (1999). Inventarisatie van de Aziatische grondeekhoorn in De Panne. Een project van de UA groep Dierenecologie in opdracht van AMINAL afdeling Natuur. Rapport 2 - Augustus-Oktober 1999.
  • Wauters, L.A. & Gurnell, J. (1999) The mechanism of replacement of red squirrels by grey squirrels: a test of the interference competition hypothesis. Ethology 105: 1053-1071.

Selected Pre-1999

  • Lurz, P.W.W., Garson, P.J. & Rushton, S.P., (1995). The ecology of squirrels in spruce dominated plantations: implications for management. Forest Ecology and Management, 79, 79-90.
  • Lurz, P.W.W., Garson, P.J., Wauters, L. (1997). Effect of temporal and spatial variation in habitat quality on red squirrel, Sciurus vulgaris, dispersal behaviour. Animal Behaviour 54, 427-435.
  • Lurz, P.W.W., Garson P.J. & Ogilvie J.F. (1998) Conifer species mixtures, cone crops and red squirrel conservation Forestry 71, 67-71.
  • Rushton, S.P., Lurz, P.W.W., Fuller, R. & Garson, P.J. (1997). Modelling the distribution of the red and grey squirrel at the landscape scale: a combined GIS and population dynamics approach. J. Appl. Ecol., 34, 1137-1154.
  • Rushton, S.P. , Lurz, P.W., Fuller, R. and Garson, P.J. (1997) Modelling the distribution of the red and grey squirrel at the landscape scale: a combined GIS and population dynamics approach. J. Appl. Ecol, 34, 1137-1154.
  • Verbeylen, G. & Matthysen, E. (1998). Inventarisatie van de Aziatische grondeekhoorn in De Panne. Een project van de UIA groep Dierenecologie in opdracht van AMINAL afdeling Natuur. Rapport September-November 1998.
  • Verbeylen, G. & Wauters, L. (1997). Habitatversnippering en eekhoorns : een vergelijking met vogelstudies. Limosa, 70, 38.
  • Villalba, S., Gulinck, H., Verbeylen, G. & Matthysen, E. (1998). Relationship between patch connectivity and the occurrence of the European red squirrel, Sciurus vulgaris, in forest fragments withi n heterogeneous landscapes. In : Dover, J.W. & Bunce, R.G.H. (eds.). Key Concepts in Landscape Ecology. IALE(UK), Preston, 205-220.

Bedtime listening

On Monday 4th August 2003, a radio programme was broadcast of BBC Radio 4 at 21.00 (British Summer Time) called, Can the red squirrel survive in Europe if the grey squirrel spreads over the Italian Alps?. To quote from the BBC website:

"Lionel Kelleway travels to Italy, to discover how the grey squirrel arrived there after World War Two and has now travelled north to the Italian Alps. In its wake it has devastated the native red squirrel population and is poised to disperse across Europe. With the help of continuous forest and few physical barriers, their spread is likely to be rapid. Lionel asks if there is hope for the red squirrel if the grey squirrel crosses Italy's border and colonises the rest of mainland Europe."

The programme features Piero Genovesi, Italo Currado, Sandro Bertolino and Luc Wauters. To listen to the programme again, go to the Radio 4 Wild Europe website: http://www.bbc.co.uk/radio4/science/wildeurope.shtml. You will need a copy of "Real One Player" to listen to the broadcast, downloadable free from the link on the BBC page.

Forestry Commission Publications

Management of grey squirrels web pages: http://www.forestresearch.gov.uk/greysquirrels

Information about FC publications in general can be found here: http://www.forestry.gov.uk/website/publications.nsf